Class CUBOZOA Werner 1975
Order CUBOMEDUSAE Haeckel 1877
The taxonomic rank and phylogenetic position of the box jellyfishes has been
much debated. They were originally considered to be an order (Cubomedusae Haeckel
1877) in the Class Scyphozoa (e.g. Mayer 1910; Kramp 1961). In the mid-1970's,
however, they were given the status of a class (Cubozoa Werner 1975; see also
Mianzan & Cornelius 1999) on the basis of morphological differences. Recent
molecular data, however, raised a paradox in that Class Cubozoa was more closely
related than Order Stauromedusae to the other scyphozoan orders Coronatae, Semaeostomeae,
and Rhizostomeae, i.e. "Scyphozoa" was paraphyletic (e.g. Collins
2002). This issue has now been resolved with elevation of stauromedusae to Class
Staurozoa (e.g. Marques & Collins 2004; Dawson in press). The list of families,
genera, and species below is based largely on Kramp (1961), Southcott (1967),
Pagès et al. (1992), and Mianzan & Cornelius (1999). The group is
currently the subject of several morphological and molecular investigations
which should help to clarify species relationships and boundaries.
Type = *
| Family (2) | Genus (8) | Species (19) | Image |
| Carybdeidae Gegenbaur 1856 | Carukia Southcott 1967 | barnesi Southcott 1967 * | |
| Carybdea Péron & Lesueur 1809 | alata Reynaud 1830 | ||
| marsupialis (Linné 1758) * | |||
| rastoni Haacke 1886 | |||
| sivickisi Stiasny 1926 | image | ||
| xaymacana Conant 1897 | |||
| Manokia Southcott 1967 | stiasnyi (Bigelow 1938) * | ||
| Tamoya Müller 1859 | haeckeli Southcott 1967 | ||
| haplonema Müller 1859 * | image | ||
| Tripedalia Conant 1897 | binata Moore 1988 | ||
| cystophora Conant 1897 * | image | ||
| Chirodropidae Haeckel 1877 | Chirodropus Haeckel 1880 | gorilla Haeckel 1880 * | |
| palmatus Haeckel 1880 | |||
| sp. Stiasny 1922 | |||
| Chironex Southcott 1956 | fleckeri Southcott 1956 * | ||
| Chiropsalmus L. Agassiz 1862 | buitendijki Horst 1907 | ||
| quadrigatus Haeckel 1880 | image | ||
| quadrumanus (Müller 1859) * | image | ||
| zygonema Haeckel 1880 |
Compiled by M.N Dawson, L.A. Gershwin, G. Jarms, H. Mianzan, A.C. Morandini.